Bo CAI,Xiang JI,Yingyong WANG,et al.An Annotated List of Lizards (Sauria: Squamata) Recorded from the People’s Republic of China[J].Asian Herpetological Research(AHR),2022,13(1):64-74.[doi:10.16373/j.cnki.ahr.210020]
Click Copy

An Annotated List of Lizards (Sauria: Squamata) Recorded from the People’s Republic of China
Share To:

Asian Herpetological Research[ISSN:2095-0357/CN:51-1735/Q]

2022 VoI.13 No.1
Research Field:
Publishing date:


An Annotated List of Lizards (Sauria: Squamata) Recorded from the People’s Republic of China
Bo CAI123 Xiang JI4 Yingyong WANG5 Dingqi RAO6 Song HUANG7 Yuezhao WANG1 Zhaobin SONG2 Xianguang GUO1* and Jianping JIANG1*
1 Chengdu Institute of Biology, Chinese Academy of Sciences, Chengdu 610041, Sichuan, China
2 Key Laboratory of Bio-resources and Eco-environment of Ministry of Education, College of Life Sciences, Sichuan University, Chengdu 610065, Sichuan, China
3 University of Chinese Academy of Sciences, Beijing 100049, China
4 Jiangsu Key Laboratory for Biodiversity and Biotechnology, College of Life Sciences, Nanjing Normal University, Nanjing 210046, Jiangsu, China
5 State Key Laboratory of Biocontrol/The Museum of Biology, School of Life Sciences, Sun Yat-sen University, Guangzhou 510275, Guangdong, China
6 Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming 650223, Yunnan, China
7 College of Life Sciences, Anhui Normal University, Wuhu 241002, Anhui, China
classification type specimen type locality distribution endemic species invasive species
From the perspectives of biodiversity conservation and management, there is an urgent need to have at hand current synopses of classification and distributions of species. In this paper, we review and summarize the classifications, Chinese and English names, type specimens, type localities and distributions of China’s lizard fauna to promote scientific exchange and species conservation among relevant people in China and internationally. As of December 31, 2020, a total of 230 species of lizard, belonging to 44 genera and 12 families, have been recorded from China, including 4 invasive species: 2 in the family Gekkonidae, 1 in Iguanidae and one in Dactyloidae. There are 115 endemic species, accounting for 50% of this group. The proportion of endemic species to China was the highest in the family Eublepharidae (84.62%), and the number of endemic species to China was the highest in the family Agamidae (42 species). The species distributions and classification in the “Fauna Sinica (Reptilia 2): Squamata (Lacertilia)” were updated. Among the provincial administrative units, there is a larger number of species in the southern provinces, such as Yunnan, Guangdong, Guangxi and Taiwan. Sichuan, Yunnan, and Taiwan harbour relatively more endemic species than others. A total of 154 species of lizards were first discovered in China, and the type localities of these species are concentrated in Yunnan, Taiwan, Tibet, Xinjiang, and Sichuan.


Agarwal I., Mahony S., Giri V. B., Chaitanya R., Bauer A. M. 2018. Two new species of bent toed geckos, Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) from Northeast India with comments on name-bearing types from the region. Zootaxa, 4420(3): 334?356
Ananjeva N. B., Guo X. G., Wang Y. Z. 2011. Taxonomic diversity of agamid lizards (Reptilia, Sauria, Acrodonta, Agamidae) from China: A comparative analysis. Asian Herpetol Res, 2(3): 117?128
Anderson J. 1872. On some Persian, Himalayan, and other Reptiles. Pro Zool Soc London, 1872: 390
Auffenberg W. 1994. The Bengal Monitor. Gainesville, USA: University Presses of Florida
Baig K. J., Philipp W., Ananjeva N. B., Wolfgang B. 2012. A morphology-based taxonomic revision of Laudakia Gray, 1845 (Squamata: Agamidae). Vertebr Zool, 62(2): 213?260
Barabanov A. V. 2009. Aspidorhinus Eichwald, 1841 as a valid subgeneric name for Eremias velox species group (Sauria, Lacertidae). Curr Stud Herpetol, 9: 59–61 (In Russian with English abstract)
Barabanov A. V., Ananjeva N. B. 2007. Catalogue of the available scientific species-group names for lizards of the genus Phrynocephalus Kaup, 1825 (Reptilia, Sauria, Agamidae). Zootaxa, 1399(1): 1?56
Bauer A. M., Rafaqat M., James T. M., Matthew P. H., Juan D. D., Todd R. J. 2013. A preliminary phylogeny of the Palearctic naked-toed geckos (Reptilia: Squamata: Gekkonidae) with taxonomic implications. Zootaxa, 3599(4): 301?324
Bedriaga J. V. 1907a. Neue Saurier aus Russisch-Asien. Annu Mus Zool Acad Impér Sci, 10(3-4): 210?243 (In German)
Bedriaga J. V. 1907b. Verzeichnis der von der Central-Asiatischen Expedition unter Stabs-Kapit?n W. Roborowski in den Jahren 1893-1895 gesammelten Reptilien. Annu Mus Zool Acad Impér Sci, 10(3-4): 181?192 (In German)
Bedriaga J. V. 1909. Amphibien und Reptilien. In Wissenschaftliche Resultate der Reisen N.M. Przewalskijs durch Zentralasien. Zool Teil Band 3 Part 1 Lacertilia Sankt-Petersburg. 3(1): 73?102, 367 (In German)
Blanford W. T. 1875. List of Reptilia and Amphibia collected by the late Dr. Stoliczka in Kashmir, Ladák, eastern Turkestán, and Wakhán, with descriptions of new species. J Asiatic Soc Bengal Calcutta, new ser, 44. Part 2(3): 191?196
Bohme W. 2003. Checklist of the living monitor lizards of the world (family Varanidae). Zool Verh, 341: 3?43
Bohme W., Ziegler T. 1997. On the synonymy and taxonomy of the Bengal monitor lizard, Varanus bengalensis (Daudin, 1802) complex (Sauria: Varanidae). Amphib Reptil, 18(2): 207?211
Cai B., Wang Y. Z., Chen Y. Y., Li J. T. 2015. A revised taxonomy for Chinese reptiles. Biodivers Sci, 23(3): 365–382 (In Chinese with English abstract)
Chaitanya R., Giri V. B., Deepak V. 2017. On the recent designation of a neotype for the taxon Calotes versicolor (Daudin, 1802) (Reptilia: Agamidae). Zootaxa, 4317(3): 585–587
Che J., Jiang K., Yan F., Zhang Y. 2020. Amphibians and reptiles in Tibet: Diversity and evolution. Beijing: Science Press (In Chinese)
Chirikova M. A., Dujsebayeva T. N., Liu J. L., Guo X. G. 2019. Geographical distribution and morphological variability of the rapid racerunner, Eremias velox (Pallas, 1771) (Reptilia, Lacertidae) in the eastern periphery of its range. Asian Herpetol Res, 10(4): 230?245
Cox C. B., Ladle R. J., Moore P. D. 2019. Biogeography: An ecological and evolutionary approach (10th edition). Hoboken, USA: Wiley
Das I., Dattagupta B. 1997. On Chinese reptile types collected during the second Forsyth expedition to Yarkand, Eastern Turkestan (1873). Sichuan J Zool, 16 (4): 168?170
Das S., Raha S., Mitra S., Bag P. 2017. Implication of variable characters for the taxonomy of Altiphylax stoliczkai (Steindachner, 1867) and Altiphylax yarkandensis (Anderson, 1872) (Reptilia: Gekkonidae) with comments on the ZSI ‘syntype’ of the latter taxon’. Zootaxa, 4320(1): 183?192
DeLisle H. F. 2009. Catalog of the genus Varanus (Reptilia: Squamata: Varanidae) with new designations of a neotype and a lectotype. ZooNova, 1(2): 8?32
Deng X. J., Ye Y. Y., Yang D. L. 1998. Two reptile records new to Hunan Province. Sichuan J Zool, 17(2): 153 (In Chinese with English abstract)
Dunayev Е. А. 1989. Taxonomic status features of ecology and behavior Phrynocephalus melanurus Eichwaid, 1831 (Reptilia: Agamidae). Bull Moscow Soc Natur Biol Dep, 94, 41–53 (In Russian)
Dunayev E. A. 2020. History of study, taxonomy, distribution, and ecology of Phrynocephalus nasatus Golubev et Dunayev, 1995 (Reptilia: Agamidae). Russ J Herpetol, 27(2): 87?96
Eremchenko V. K., Panfilov, A. M. 1999. Taxonomic situation of multiocellated racerunner of the Eremias multiocellata complex of Kyrgyzstan and neighbour China (Sauria: Lacertidae: Eremias). Sci New Technol, 4: 112?124 (In Russian)
Eremchenko V. K., Panfilov, A. M., Tzarinenko, E. I. 1992. Eremias multiocellata complex: Solution of some problems in systematics of the multiocellated racerunners of Kyrgyzstan (Sauria, Lacertidae, Eremias) 65?80. In Eremchenko V. K., Panfilov, A. M., Tzarinenko, E. I. Conspectus of the Researches on Cytogenetics and Systematics of Some Asiatic Species of Scincidae and Lacertidae. Bishkek, Ilim Publishing (In Russian)
Favre A., P?ckert M., Pauls S. U., J?hnig S. C., Uhl D., Michalak I., Muellner-Riehl A. N. 2015. The role of the uplift of the Qinghai-Tibetan Plateau for the evolution of Tibetan biotas. Biol Rev, 90(1): 236?253
Freitas E. S., Datta-Roy A., Karanth P., Grismer L. L., Siler C. D. 2019. Multilocus phylogeny and a new classification for African, Asian and Indian supple and writhing skinks (Scincidae: Lygosominae). Zool J Linn Soc, 186(4): 1067–1096?
GBIF Secretariat. 2020. GBIF backbone taxonomy. Retrieved from, 30/12/2020
Giri V. B., Chaitanya R., Mahony S., Lalrounga S., Lalrinchhana C., Das A., Sarkar V., Karanth P., Deepak V. 2019. On the systematic status of the genus Oriocalotes Günther, 1864 (Squamata: Agamidae: Draconinae) with the description of a new species from Mizoram state, Northeast India. Zootaxa, 4638(4): 451–484
Gowande G., Mishra A., Mirza Z. A. 2016. Neotype designation for Calotes versicolor Daudin, 1802 (Sauria: Agamidae) with notes on its systematics. Zootaxa, 4126(2): 271–279
Gowande G. G., Bhosale H. S., Phansalkar P. U., Sawant M., Mirza Z. A. 2021. On the systematics and the phylogenetic position of the poorly known, montane dragon-lizard species Pseudocalotes austeniana (Annandale, 1908) (Squamata, Agamidae, Draconinae). Evol Syst, 5(1): 141–150
Grismer L. L., Ngo H. N., Qi S., Wang Y. Y., Le M. D., Ziegler T. 2021. Phylogeny and evolution of habitat preference in Goniurosaurus (Squamata: Eublepharidae) and their correlation with karst and granite-stream-adapted ecomorphologies in species groups from Vietnam. Vertebr Zool, 71: 335–352
Grismer L. L., Shi H. T., Nikolai L. O., Natalia B. A. 2002. A new species of Goniurosaurus (Squamata: Eublepharidae) from Hainan Island, China. J Herpetol, 36(2): 217?224
Guo X. G., Chen D. L., Wan H. F., Wang Y. Z. 2010. Review of systematics on the racerunner lizard (Lacertidae: Eremias). Sichuan J Zool, 29(4): 665?672 (In Chinese with English abstract)
Hawkins B. A., Field R., Cornell H. V., Currie D. J., Guegan J. F., Kaufman D. M., Kerr J. T., Mittelbach G. G., Oberdorff T., O’Brien E. M. 2003. Energy, water, and broad-scale geographic patterns of species richness. Ecology, 84 (12): 3105?3117
Herpetological Department. 1977. Retri of Chinese reptile taxonomy. Beijing, China: Science Press (In Chinese)
Hsü K. J., Chen H. 1999. Geological atlas of China: An application of the tectonic facies concept to the geology of China. New York, USA: Elsevier Science.
Huang Y., Dai D., Chen Y. Y., Wan H. F., Li J. T., Wang Y. Z. 2011. Lizard species richness patterns in China and its environmental associations. Biodivers Conserv, 20: 1399–1414
Hsiang K. S., Lee P. X., Yang Y. R. 2009. Atlas of amphibians and reptiles in Taiwan. Taipei, China: Owl Publishing House (In Chinese)
ICZN (International Comission on Zoological Nomenclature). 1999. International Code of zoological nomenclature. Fourth Edition adopted by the International Union of Biological Sciences. London, UK: International Trust of Zoological Nomenclature
Jin Y. T., Brown R. P. 2013. Species history and divergence times of viviparous and oviparous Chinese toad-headed sand lizards (Phrynocephalus) on the Qinghai-Tibetan Plateau. Mol Phylogenet Evol, 68(2): 259?268
Jin Y. T., Brown R. P. 2019. Morphological species and discordant mtDNA: A genomic analysis of Phrynocephalus lizard lineages on the Qinghai-Tibetan Plateau. Mol Phylogenet Evol, 139: 106523
Jin Y. T., Yang Z. S., Brown R. P., Liao P., Liu N. 2014. Intraspecific lineages of the lizard Phrynocephalus putjatia from the Qinghai-Tibetan Plateau: Impact of physical events on divergence and discordance between morphology and molecular markers. Mol Phylogenet Evol, 71: 288?297
Kimmins J. P. 1987. Forest ecology. New York, USA: Macmillan
Li P. P., Zhao E. M., Dong B. J. 2010. Amphibians and reptiles of Tibet. Beijing, China: Science Press (In Chinese)
Lin J. T., Yao C. J. 2016. A new species of the genus Gekko (Squamata: Sauria: Gekkonidae) from Guishan Isle, Yilan County, Taiwan. J Nat Taiwan Mus, 69(2): 1?14 (In Chinese with English abstract)
Liu J. L. 2019. Molecular phylogeny and phylogeography of Eremias velox complex and Eremias multiocellata complex. Ph.D Thesis. Sichuan University. 179 pp (In Chinese with English abstract)
Liu J. L., Guo X. G., Chen D. L., Li J., Yue B. S., Zeng X. M. 2019. Diversification and historical demography of the rapid racerunner (Eremias velox) in relation to geological history and Pleistocene climatic oscillations in arid Central Asia. Mol Phylogenet Evol, 130: 244?258
Liu S., Hou M., Wang J., Ananjeva N. B., Rao D. Q. 2020. A new species of Diploderma (Squamata: Sauria: Agamidae) from Yunnan Province, China. Russ J Herpetol, 27(3): 127?148
Liu S., Rao D. Q. 2019. A new species of the genus Acanthosaura from Yunnan, China (Squamata, Agamidae). ZooKeys, 888(3): 105?132
Mahony S. 2010. Systematic and taxomonic ruation of four little known Asian agamid species, Calotes kingdonwardi Smith, 1935, Japalura kaulbacki Smith, 1937, Salea kakhienensis Anderson, 1879 and the monotypic genus Mictopholis Smith, 1935 (Reptilia: Agamidae). Zootaxa, 2514: 1?23
Meng K., Li S., Murphy R. W. 2008. Biogeographical patterns of Chinese spiders (Arachnida: Araneae) based on a parsimony analysis of endemicity. J Biogeogr, 35(7): 1241?1249
Melville R. V. 1981. Eremias Wiegmann, 1834 (Reptilia Lacertilia): Proposed designation of a type species by use of the plenary powers. Bull Zooll Nomencl, 38: 283-284
Melville R. V. 1985. Opion 1318 Lacerta velox Pallas, 1771 is the type species of Eremias Wiegmann , 1834. Bull Zooll Nomencl, 42(2): 171?172
Melville J. H. J., Mantziou G., Ananjeva N. B, Milto K. D., Clemann N. 2009. Historical biogeography, phylogenetic relationships and intraspecific diversity of agamid lizards in the Central Asian deserts of Kazakhstan and Uzbekistan. Mol Phylogenet Evol, 53(1): 99?112
Milto K. D., Barabanov A. V. 2012. A catalogue of the agamid and chamaeleonid types in the collection of the Zoological Institute, Russian Academy of Sciences, St. Petersburg. Russ J Herpetol, 19(2): 155?170
Ngo H. N., Ziegler T., Nguyen T. Q., Pham C. T., Nguyen T. T., Le M. D., van Schingen M. 2016. First population assessment of two cryptic Tiger Geckos (Goniurosaurus) from northern Vietnam: Implications for conservation. Amphib Reptile Conse, 10(1): 34-45
Orlova V. F., Poyarkov N. A., Chirikova M. A., Nazarov R. A. 2017. MtDNA differentiation and taxonomy of Central Asian racerunners of Eremias multiocellata-E. przewalskii species complex (Squamata, Lacertidae). Zootaxa, 4282(1): 1?42
Ota H. 2000. Japalura szechwanensis, a junior synonym of J. fasciata. J Herpetol, 34(4): 611?614
Peng L. F. 2019. Species diversity and phylogeography of the genus Laudakia (Iguania; Agamidae) in China. Ph.D Thesis. Tibet University. 134 pp (In Chinese with English abstract)
Pianka E. R., King D. R. 2004. Varanoid lizards of the world. St, Bloomington, USA: Indiana University Press
Pope C. H. 1935. The reptiles of China. Natural History of Central Asia, Vol.10. New York: USA, American Museum of Natural History
Pyron R. A., Burbrink F. T., Wiens J. J. 2013. A phylogeny and revised classification of Squamata, including 4161 species of lizards and snakes. BMC Evol Biol, 13: 93
Rao D. Q., Yang D. T. 1996. The preliminary researches on the varanid species of Yunnan and their habitats, habits and conservation situation. Biodivers Sci, 4(4): 194?200 (In Chinese with English abstract)
Renner S. S. 2016. Available data point to a 4-km-high Tibetan Plateau by 40 Ma, but 100 molecular-clock papers have linked supposed recent uplift to young node ages. J Biogeogr, 43(8): 1479–1487
Rosenzweig M. L. 1995. Species diversity in space and time. England, UK: Cambridge Unievrsity Press
Rosler H. 2017. Gecko-Chorologie (Squamata: Gekkota). Gekkota, (4): 1–160
Schmidtler J. F., B?hme W. 2011.?Synonymy and nomenclatural history of the common or viviparous lizard, by this time:?Zootoca vivipara?(Lichtenstein, 1823).?Bonn Zool Bull,?60(2): 214–228
Schneider J. G. 1792. Amphibiorum physiologiae specimen alterum historiam et species generis stellionum seu geckonum sistens. Traiecti ad Viadrum, C. L. F. Aitzi, 2: 30?31 (In Latin)
Schneider J. G. 1797. Amphibiorum physiologiae specimen alterum historiam et species generis stellionum seu geckonum sistens. Züllichoviea, apud Friedericum Frommannum, 30?31 (In Latin)
Shen Y. H., Ye Y. Y., Deng X. J. 2014. Fauna Hunan: Reptilia. Hunan, China. Changsha, China: Hunan Science and Technology Press (In Chinese)
Shi L., Lin X. L., Zhao H. 2011. New record of Hemiphyllodactylus yunnanensis to Xizang Autonomous Region, China. Chin J Zool, 46(1): 131?133 (In Chinese with English abstract)
Sindaco R., Eremchenko V. K. 2008. The reptiles of the Western Palearctic. 1. Annotated checklist and distributional atlas of the turtles, crocodiles, amphisbaenians and lizards of Europe, north Africa, Middle East and Central Asia. Latina, Italy: Edizioni Belvedere
Smith M. A. 1935. The fauna of British India, including Ceylon and Burma. Reptilia and Amphibia. II. Sauria. London, UK: Taylor & Francis Ltd
Solovyeva E. N., Lebedev V. S., Dunayev E. A., Nazarov R. A., Bannikova A. A., Che J., Murphy R. W., Poyarkov N. A. 2018. Cenozoic aridization in Central Eurasia shaped diversification of toad-headed agamas (Phrynocephalus, Agamidae, Reptilia). PeerJ, 6: e4543
Solovyeva E. N., Poyarkov N. A., Dunaev E. A., Duysebayeva T. N., Bannikova A. A. 2011. Molecular differentiation and taxonomy of the sunwatcher toad-headed agama species complex Phrynocephalus superspecies helioscopus (Pallas 1771) (Reptilia: Agamidae). Russell J Genet, 47(7): 842?856
Song M. T. 1985. A new species of Gekko from Shaanxi. Acta Herpetol Sin, 4(4): 329–330 (In Chinese with English abstract)
Steindachner F. 1870. Herpetologische Notizen (II). Reptilien gesammelt W?hrend einer Reise in Sengambien. Sitzungsber Kais Akad Wiss Wien, 62: 326?349 (In German)
Sunandan D., Raha S., Mitra S., Bag P. 2017. Implication of variable characters for the taxonomy of Altiphylax stoliczkai (Steindachner, 1867) and Altiphylax yarkandensis (Anderson, 1872) (Reptilia: Gekkonidae) with comments on the ZSI ‘syntype’ of the latter taxon. Zootaxa, 4320 (1): 183–192
Szczerbak N. N., Golubev M. L. 1977. Systematics of the Palearctic geckos (genera Gymnodactylus, Bunopus, Alsophylax). Proc Zool Inst Acad Sci USSR, 74: 120–133 (In Russian)
Uetz P., Freed P., Ho?ek J. 2020. The reptile database. Retrieved from 30/12/2020
Wan T. 2003. The outline of geotectonics of China. Beijing, China: Geological Publishing House (In Chinese)
Wang K., Ren J., Chen H., Lyu Z., Guo X., Jiang K., Chen J., Li J., Guo P., Wang Y., Che J. 2020. The updated checklists of amphibians and reptiles of China. Biodivers Sci, 28 (2): 189–218 (In Chinese with English abstract)
Wang K., Che J., Lin S., Deepak V., Aniruddha D., Jiang K., Jin J., Chen H., Siler C. D. 2019. Multi-locus phylogeny and taxonomic revision of the Mountain Dragons of the genus?Japaluras. l.?Gray, 1853 (Reptilia: Squamata: Agamidae). Zool J Linn Soc, 185: 246–267
Wang K., Jiang K., Deepak V., Abhijit D., Hou M., Che J., Siler C. D. 2018. On the occurrences of Japalura kumaonensis and Japalura tricarinata (Reptilia: Sauria: Draconinae) in China. Herpetologica, 74(2): 181?190
Wang Y. Y., Liu Y. 2014. Atlas of terrestrial vertebrates in Heishiding. Beijing, China: Higher Education Press (In Chinese with English abstract)
Wood Jr. P. L., Guo X. G., Travers S. L., Su Y. C., Olson K. V., Bauer A. M., Grismer L. L., Siler C. D., Moyle R. G., Andersen M. J., Brown R. M. 2020. Parachute geckos free fall into synonymy: Gekko phylogeny, and a new subgeneric classification, inferred from thousands of ultraconserved elements. Mol Phylogenet Evol, 146: 106731
Yang D. T., Rao D. Q. 2008. Amphibia and Reptilia of Yunnan. Kunming, China: Yunnan Science and Technology Press (In Chinese)
Yang J. H., Wang H. C., Ma X. X., Wang Y. Y. 2009. New distribution records of Sphenomorphus maculatus and additional description of the species. Chin J Zool, 44(6): 156?159 (In Chinese with English abstract)
Yu D. N. 2014. The study on phylogenetic relationship and interspecific hybridization of Takydromus in China. Ph.D Thesis. Nanjing Normal University. 141 pp (In Chinese)
Yuan M., Li D. H., Zhang L. 2009. Scincella potanini: A new genus and a new species record to Guizhou. Sichuan J Zool, 28(3): 427+443+481 (In Chinese with English abstract)
Zhang B., Qian T., Jiang X., Cai B., Deng X., Yang D. 2020. A new species of Hemiphyllodactylus Bleeker, 1860 (Reptilia: Squamata) from Hunan Province, China. Asian Herpetol Res, 11(3): 183–193
Zhang J. J. 2019. Diversity and protection of amphibians and reptiles in Yangcheng Manghe Rhesus Monkey National Nature Reserve, Yangcheng, Shanxi Province. ChinJ Wildl, 40(4): 969?966 (In Chinese with English abstract)
Zhang R. Z. 2011. Zoogeography of China. Beijing, China: Science Press (In Chinese)
Zhang Y. Y., Chen C. T., Li L., Zhao C. J., Chen W. C., Huang Y. 2014. Insights from ecological niche modeling on the taxonomic distinction and niche differentiation between the black-spotted and red-spotted tokay geckoes (Gekko gecko). Ecol Evol, 4(17): 3383?3394
Zhao E. M., 1997. Notes on amphibians and reptiles of Hong Kong. Sichuan J Zool, 16(2): 51?60 (In Chinese with English abstract)
Zhao E. M. 2003. Coloured atlas of reptiles of Sichuan. Beijing, China: China Forestry Publishing House (In Chinese)
Zhao E. M., Adler K. 1993. Herpetology of China. Oxford, UK: Society for the Study of Amphibians and Reptiles
Zhao E. M., Zhao H., Zhou Z. Y. 2004. Herpetodiversity of northeastern China and their distribution. Sichuan J Zool, 23(3): 165?168 (In Chinese with English abstract)
Zhao E. M., Zhao K. T., Zhou K. Y. 1999. Fauna Sinica, Reptilia, vol 2: Squamata, Lacertilia. Beijing, China: Science Press (In Chinese)
Zhao E. M., Zhang X. W., Zhao H., Adler K. 2000. Revised checklist of Chinese Amphibia and Reptilia. Sichuan J Zool, 19(3): 196?207 (In Chinese with English abstract)
Zhao K. T. 1995. The Eremias quadrifrons (Strauch) and Pelochelys taihuensis Zhang are thought to be invalid species. J Shuzhou Railway Teach Coll (Nat Sci), 12(2): 31?35 (In Chinese with English abstract)
Zhou K. Y., Liu Y. Z., Yang G. P. 1981. Three new subspecies of Hemiphyllodactylus yunnanensis (Boulenger) (Lacertiformes: Gekkonidae) from China. Acta Zootax Sin, 6(2): 202?210 (In Chinese with English abstract)


Last Update: 2022-03-25