Barbod SAFAEI-MAHROO,Hanyeh GHAFFARI,Hadi FAHIMI,et al.The Herpetofauna of Iran: Checklist of Taxonomy, Distribution and Conservation Status[J].Asian Herpetological Research(AHR),2015,6(4):257-290.[doi:10.16373/j.cnki.ahr.140062]
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The Herpetofauna of Iran: Checklist of Taxonomy, Distribution and Conservation Status
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Asian Herpetological Research[ISSN:2095-0357/CN:51-1735/Q]

2015 VoI.6 No.4
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The Herpetofauna of Iran: Checklist of Taxonomy, Distribution and Conservation Status
1 Pars Herpetologists Institute, 3th Floor, No. 24, Roodbar S.W alley, Mirdamad, Tehran, Iran
2 Department of Environmental Sciences, Faculty of Natural Resources, University of Kurdistan, Sanandaj, Iran
3 Department of Biology Faculty of Science Ege University, ?zmir, Turkey
4 Department of Biology, Faculty of Science, Ferdowsi University of Mashhad, Mashhad, Iran
5 Department of Biology, Faculty of Scince, Razi University, Kermanshah, Iran
Amphibians Anura Caudata Reptiles Crocodylia Squamata Testudines Persian common name
We present an annotated checklist for a total 241 reptiles and 22 amphibians including 5 frogs, 9 toads, 7 newts and salamanders, 1 crocodile, 1 worm lizard, 148 lizards, 79 snakes and 12 turtles and tortoises, includes the most scientific literature up to August 2014 and also based on several field surveys conducted in different Provinces of Iran from 2009 to 2014. We present an up-to-dated checklist of reptiles and amphibians in Iran. We provide a comprehensive listing of taxonomy, names, distribution and conservation status of all amphibians and reptiles of Iran. This checklist includes all recognized named taxa, English names for classes, orders, families, species, subspecies along with Persian names for species, including indication of native and introduced species. For the first time we report two non-native introduced reptiles from natural habitats of Iran. Of the total 22 species of amphibians in Iran, 6 (27.2%) are endemic and of the total 241 species of reptiles, 55 (22.8%) are endemic. Of the 22 amphibians species in Iran, 3 (13%) are Critically Endangered, 2 (9%) are Vulnerable and of the 241 reptile species 3 (1.2%) are Critically Endangered, 4 (1.6%) are Endangered and 10 (4.1%) are Vulnerable. Accordingly, this paper combines significant aspects of taxonomy, common names, conservation status and distribution of the Iranian herpetofauna.


Abtin E., Nilson G., Mobaraki A., Hosseini A. A., Dehgannejhad M. 2014. A New Species of Krait, Bungarus (Reptilia, Elapidae, Bungarinae) and the First Record of That Genus in Iran. Russ J Herpetol, 21: 243–250
Ahmadzadeh F., Flecks M., Torki F., B?hme W. 2011. A new species of angular-toed gecko, genus Cyrtopodion (Squamata: Gekkonidae), from southern Iran. Zootaxa, 2924: 22–32
Ahmadzadeh F., Flecks M., Carretero M. A., Mozaffari O., B?hme W., Harris D. J., Freitas S., R?dder D. 2013. Cryptic Speciation Patterns in Iranian Rock Lizards Uncovered by Integrative Taxonomy. Plos One, 8(12): e80563
Ananjeva N. B., Orlov N. L., Khalikov R. G., Darevsky I. S., Ryabov S. A., Barabanov A. V. 2006. The Reptiles of Northern Eurasia: Taxonomic Diversity, Distribution, Conservation Status. Zool Inst Russ Acad Sci, Pensoft Series Faunistica, No, 47, Sofia. 246 pp
Anderson J. 1872. On some Persian, Himalayan, and other reptiles. Proc Zool Soc, XXV: 371–404
Anderson S. C. 1970. Checklist and key to the lizards of Iran. Pages 284-285 in Year Book of the American Philosophical Society, vol.1969. American Philosophical Society, Philadelphia, Pennsylvania, USA
Anderson S. C. 1974. Preliminary key to the turtles, lizards and amphisbaenians of Iran. Fieldiana Zool, 65: 27–44
Anderson S. C. 1999. The Lizards of Iran. New York, USA: SSAR Ithaca, 442 pp
Baig J. K., Masroor R. 2008. The snakes of the genus Spalerosophis JAN, 1865 in Indo-Pakistan and Iran. Herpetozoa, 20: 109–115
Baloutch M., Kami H. G. 1995. Amphibians of Iran. Tehran, Iran: University of Tehran Press, 177 pp
Beolens B., Watkins M., Grayson M. 2011. The Eponym Dictionary of Reptiles. Baltimore, USA: The Johns Hopkins University Press, xiii+296 pp
Beolens B., Watkins M., Grayson M. 2013. The Eponym Dictionary of Amphibians. Exeter, UK: The Johns Hopkins University Press, xiii+244 pp
Blanford W. T. 1876. The Zoology and Geology. In William T. Blanford, ed., Eastern Persia, an Account of the Journeys of the Persian Boundary Commission, 1870–1872 vol. 2. London, England, UK: Macmillan and Co., 516 pp
Bostanchi H., Anderson S. C., Kami H. G., Papenfuss T. J. 2006. A new species of Pseudocerastes with elaborate tail ornamentation from western Iran (Squamata: Viperidae). Proc Calif Acad Sci ser, 4: 443-450
Boulenger G. A. 1899. Reptilia and Amphibia of Lake Urmi and its neighborhood. J Proc Linn Soc, 27: 378–381
Boulenger G. A. 1908. Description of a new species of Lacerta from Persia. Proc Zool Soc London, 1908: 934–936
Boulenger G. A. 1920. Description of a new snake of the genus Contia, B. & G., from Persia. J Bombay nat hist Soc, 26: 1024
Boulenger G. A. 1921. Description of a new Land-Tortoise from Northern Persia. J Bombay nat hist Soc, 27: 251–252
Brandt M. 1837. Note sur quatre nouvelles espèces de serpents de la c?te occidentale de la mer Caspienne et de la Perse septentrionale, decouvertes par M. Kareline. Bull Sci Acad Imp Sci St. Petersb, III: 241–244
Carranza S., Arnold E. N., Pleguezuelos J. M. 2006. Phylogeny, biogeography, and evolution of two Mediterranean snakes, Malpolon monspessulanus and Hemorrhois hippocrepis (Squamata, Colubridae), using mtDNA sequences. Mol Phylogenet Evol, 40: 532–546.
Convention on International Trade in Endangered Species of Wild Fauna and Flora. 2014. Available from (Accessed on 24 April 2014)
De Filippi F. 1863. Nuove o poco note specie di animali vertebrati raccolte in un viaggio in Persia nell’estate dell’anno 1862. Arch zool anat fisiol Modena, 2: 377–394
De Filippi F. 1864. Riassunto Del Catalogo degli Animali vertebrati delle Provincie caucasiche e della Persia occidentale. Atti Soc Ital Sci Nat, 1864: 184–186
Dubois A. Bour R. 2010. The nomenclatural status of the nomina of amphibians and reptiles created by Garsault (1764), with a parsimonious solution to an old nomenclatural problem regarding the genus Bufo (Amphibia, Anura), comments on the taxonomy of this genus, and comments on some nomina created by Laurenti (1768). Zootaxa, 2447: 1–52
Dubois A. Raffa?lli J. 2012. A new ergotaxonomy of the order Urodela Duméril, 1805 (Amphibia, Batrachia). Alytes. 28: 77–161
Eskandarzadeh N., Darvish J., Rastegar-Pouyani E., Ghassemzadeh F. 2013. Reevaluation of the taxonomic status of sand boas of the genus Eryx (Daudin, 1803) (Serpentes Boidae) in northeastern Iran. Turk J Zool, 37: 348–356
Fathinia B., Karamiani R., Darvishnia H., Heidari N., Rastegar-Pouyani N. 2011. A new species of Carinatogecko (Sauria: Gekkonidae) from Ilam Province, western Iran. Amphib reptile conserv, 5: 61–74
Fathinia B., Rastegar-Pouyani N., Rastegar-Pouyani E., Toodeh-Dehghan F., Rajabizadeh M. 2014. Molecular systematics of the genus Pseudocerastes (Ophidia: Viperidae) based on the mitochondrial cytochrome b gene. Turk J Zool, 38: doi: 10.3906/zoo–1308–25
Firouz E. 2005. The Complete Fauna of Iran. Cambridge, UK: Camb Pub Manag Ltd, 327 pp
Frank, N., Ramus E. 1995. Complete Guide to Scientific and Common Names of Amphibians and Reptiles of the World. Pottsville, Pennsylvania: N.G. Publishers. 377 pp
Fritz U., Ayaz D. Buschbom J., Kami H. G., Mazanaeva L. F., Aloufi A. A., Auer M., Rifai L., ?ili? T., Hundsd?rfer A. K. 2007.Go east: phylogeographies of Mauremys caspica and M. rivulata discordance of morphology, mitochondrial and nuclear genomic markers and rare hybridization. J Evol Biol, doi:10.1111/j.1420–9101.2007.01485.x
Frost D. R. Grant T. Faivovich J. Bain R. H. Haas A. Haddad C. F. B. De SáR. O. Channing A. Wilkinson M. Donnellan S. C. Raxworthy C. J. Campbell J. A. Blotto B. L. Moler P. Drewes R. C. Nussbaum R. A. Lynch J. D. Green D. M. Wheeler W. C. 2006. The amphibian tree of life. Bull Am Mus Nat Hist, New York, USA. 370 pp
Frynta D., Moravec J., ?ihákoá J., Sádlo J., Hodková Z., Kaftan M., Kodym P., Král D., Pitule V., ?ejna L. 1997. Results of the Czech Biological Expedition to Iran.Part 1. Notes on the distribution of amphibians and reptiles. Acta Soc Zool Bohem, 61: 3–17
Gholamhosseini A., Schmidtler J. F., Zareian H., Esmaeili H. R. 2009. The second specimen of the Iranian Eirenis rechingeri Eiselt, 1971. Herpetozoa, 22: 189–190
Gholamhosseini A., Esmaeili H. R. 2010. First record and range extension of Field’s horned viper, Pseudocerastes fieldi Schmidt, 1930 (Squamata Viperidae), from Fars province, southern Iran. Turk J Zool, 34: 551–552
Gholamifard A., Rastegar-Pouyani N., Esmaeili H. R. 2012. Annotated checklist of reptiles of Fars Province, southern Iran. IJAB, 8: 155–167
Heidari N., Rastegar-Pouyani N., Rastegar-Pouyani E., Rajabizadeh M. 2013. A new species of Acanthodactylus Fitzinger 1834 (Sauria: Lacertidae) from southern Iran. Zootaxa, 3722: 333–346
International Union For Conservation Of Nature. 2014. IUCN Red List of Threatened Species. Version 2014. 3. Available from (Accessed on 15 November 2014)
Kazemi S. M., Farhadi Qomi M., Kami H. G., Anderson S. C. 2011. A new species of Ophiomorus (Squamata: Scincidae) from Maranjab Desert, Isfahan Province, Iran, with a revised key to the genus. Amphib reptile conserv, 5: 23–33
Krause V., Ahmadzadeh F., Moazeni M., Wagner P., Wilms T. M. 2013. A new species of the genus Tropiocolotes Peters, 1880 from western Iran (Squamata: Sauria: Gekkonidae). Zootaxa, 3716: 022–038
Lantz L. A. 1918. Reptiles from the River Tajan (Transcaspia). Proc zool London, II: 11–19
Latifi M., Hoge A. R., Eliazan M. 1966. The poisonous snakes of Iran. Mem Inst Butantan, 33: 735–743
Latifi M. 1991. The Snakes of Iran, English edition. Contributions to Herpetology, vol. 7. Ohio, USA: SSAR Oxford, 159 pp
Linnaeus C. 1758. Systema Naturae per Regna Tria Naturae, Secundum Classes, Ordines, Genera, Species, cum Characteribus, Differentiis, Synonymis, Locis, 10th edition, vol. 1, part 1. Laurenti Salvii, Stockholm, Sweden. (4) + 823 + (1) pp
Litvinchuk S. N., Borkin L. J., Skorinov D.V., Rosanov J. M. 2008. A new species of common toads from the talysh mountains, south-eastern Caucasus: genome size, allozyme, and morphological evidences. Russ J Herpetol, 15: 19–43
Mahlow K., Tillack F., Schmidtler J. F., Müller J. 2013.An annotated checklist, description and key to the dwarf snakes of the genus Eirenis Jan, 1863 (Reptilia Squamata Colubridae), with special emphasis on the dentition. Vert Zool, 63: 41–85
Mashkaryan V., Vamberger M., Arakelyan M., Hezaveh N., Carretero M. A., Corti C., Harris D., J. Fritz U. 2013. Gene flow among deeply divergent mtDNA lineages of Testudo graeca (Linnaeus, 1758) in Transcaucasia. Amphibia-Reptilia, 34: 337–351
Melnikov D., Melnikova E., Nazarov R., Rajabizadeh M. 2013. Taxonomic revision of Phrynocephalus persicus DE FILIPPI, 1863 complex with description of a new species from zagros, southern Iran. СОВРЕМЕННАЯ ГЕРПЕТОЛОГИЯ. 13: 34–46
Melnikov D., Melnikova E., Nazarov R., Rajabizadeh M., Al-Johany A., Amr Z. S., Ananjeva N. B. 2014. Taxonomic revision of Phrynocephalus arabicus ANDERSON, 1984 complex with description of a new species from Ahvaz, south-western Iran. Russ J Herpetol, 21: 149–159
Mertens R. F. W. 1956. Amphibien und Reptilien aus SO-Iran. Jahrh. Jahresh Ver vaterl Naturkd Wb, 111: 90–97 (In German)
Mertens R. F. W. 1957. Weitere Unterlagen zur Herpetofauna von Iran. Jahresber. Jahresh Ver vaterl Naturkd Wb, 112: 118–128 (In German)
Murray J. A. 1884. Additions to the present knowledge of the Vertebrate Zoology of Persia. Ann Mag Nat Hist, 5: 97–106
Mozaffari O., Ahmadzadeh F., Parham J. F. 2011. Eremias papenfussi sp. nov., a new lacertid lizard (Sauria: Lacertidae) from Tehran Province, Iran. Zootaxa, 3114: 57–62
Mozaffari O., Parham J. F. 2007. A new species of racerunner lizard (Lacertidae: Eremias) from Iran. Proc Calif Acad Sci ser, 4, 58: 569–574
Nazarov R. A., Ananjeva N. B., Radjabizadeh M. 2010. Two new species of angular-toed geckos (Squamata: Gekkonidae) from southern Iran. Russ J Herpetol, 16: 311–324
Nazarov R. A., Bondarenko D. A., Radjabizadeh M. 2012. A new species of thin-toed geckos Cyrtopodion sensulato (Squamata: Sauria: Gekkonidae) from Hormozgan Province south Iran. Russ J Herpetol, 19: 292–298
Nazarov R. A., Rajabizadeh M. 2007. A new species of angular-toed gecko of the genus Cyrtopodion (Squamata: Sauria: Gekkonidae) from southeast Iran (Sistan-Baluchistan Province). Russ J Herpetol, 14: 137–144
Nikolsky A. M. 1886. Contribution to the knowledge of the fauna of vertebrate animals of northeastern Persia and Transcaucasia. Trav St Pétersbourg Soc Nat, 17: 403–406 (In Russian)
Nikolsky A. M. 1896. Diagnoses Reptiliurn et Amphibiorum novorum in Persia oriental a N. Zarudny collectorum. Ann Mus Zool Acad Imp Sci St. Pétersbourg, 4: 369–372 (In Latin)
Nikolsky A. M. 1897. Les reptiles, amphibies, et poissons, recueillis par Mr. N. A. Zarudny dans la Perse orientale. Ann Mus Zool Acad Imp Sci St. Pétersbourg, 2: 306–348 (In Russian)
Nikolsky A. M. 1899. Reptiles, amphibies et poissons recueillis pedant le voyage de Mr. N. A. Zarudny en 1898 dans la Perse. Ann Mus Zool Acad Imp Sci St. Pétersbourg, 4: 375–417 (In Russian)
Nikolsky A. M. 1903. On the species of Reptiles collected by Mr. N. Zarudny in Eastern Persia in 1901. Ann Mus Zool Acad Imp Sci St. Pétersbourg, 8: 95–98 (In Russian)
Nikolsky A. M. 1907. Presmykaiushchiiasia i zemnovoddnyia, sobrannyia N. A. Zarudym v Persii v 1903–1904 gg. Reptiles et amphibies recueillis par Mr. N. A. Zaroudny en Perse en 1903–1904. Ann Mus Zool Acad Imp Sci St. Pétersbourg, 10: 260–301 (In Russian).
Nilson G., Rastegar-Pouyani N., Rastegar-Pouyani E., Andrén C. 2003. Lacertas of south and central Zagros Mountains, Iran, with description of two new taxa. Russ J Herpetol, 10: 11–24
Olivier G. A. 1804. Voyage dans l’Empire Othoman, l’?gypte et la Perse, Fait par ordre du Gouvernement, pendant les six premières années de la République, vol. 4. Paris, France: H Agasse, 501 pp
Olson D. M., Dinerstein E., Wikramanayake E. D., Burgess N. D., Powell G. V. N, Underwood E. C., D’Amico J. A., Itoua I., Strand H. E., Morrison J. C., Loucks C. J., Allnutt T. F., Ricketts T. H., Kura Y., Lamoreux J. F., Wettengel W. W., Hedao P., Kassem K. R. 2001. Terrestrial ecoregions of the world: a new map of life on Earth. Biosci, 51: 933–938
Parham J. F., Türkozan O., Stuart B. L., Arakelyan M., Shafei S., Macey J. R., Werner Y. L., Papenfuss T. J. 2006. Genetic Evidence for Premature Taxonomic Inflation in Middle Eastern Tortoises. Proc Calif Acad Sci, 57: 955–964
Pook C. E., Joger U., Stümpel N., Wüster W. 2009. When continents collide: Phylogeny, historical biogeography and systematics of the medically important viper genus Echis (Squamata: Serpentes: Viperidae). Mol Phylogenet Evol, 53: 792–807
Procter J. B. 1921. Further Lizards and Snakes from Persia and Mesopotamia. J Bombay Nat Hist Soc, 28: 251–253
Rajabizadeh M., Nilson G., Kami H. G. 2011. A new species of mountain viper (Ophidia: Viperidae) from the central Zagros Mountains, Iran. Russ J Herpetol, 18: 235–240
Rajabizadeh M., Schmidtler J. F., Orlov N. L., Soleimani G. 2012. Review of taxonomy and distribution of the Eirenis medus group (Chernov, 1940) (Ophidia: Colubridae) with description of a new species of the genus Eirenis from Kerman Province, southeastern Iran. Russ J Herpetol, 19: 307–313
Rastegar-Pouyani N., Kami H. G., Rajabzadeh M., Shafiei S., Anderson S. C. 2008. Annotated Checklist of Amphibians and Reptiles of Iran. IJAB, 4: 43–66
Rastegar-Pouyani N., Rastegar-Pouyani E. 2001. A new species of Eremias (Sauria: Lacertidae) from highlands of Kermanshah Province, western Iran. Asiat herpetol res, 9: 107–112
Rastegar-Pouyani N., Nilson G., Faizi H. 2006. A new species of Asaccus (Sauria: Gekkonidae) from Kurdistan Province, western Iran. Hamadryad 30: 141–150
Sabeti H. 1997. Forest, Trees and Shrubs of Iran. Yazd Iran: Yazd Uni Pub, 806 pp
Sadeghi R., Torki F. 2011. Notes on the natural history and distribution of Carinatogecko stevenandersoni Torki, 2011. Amphib reptile conserv, 5: 34–36
Sch?tti B., Tillack F. Helfenberger N. 2009. A contribution to Spalerosophis microlepis JAN, 1865, with a short review of the genus and a key to the species. Herpetozoa, 22: 115–135
Schmidt K. P. 1939. Reptiles and amphibians from southwestern Asia. Zool Ser Field Mus Nat Hist, 24: 49–91
Schmidt K. P. 1955. Amphibians and Reptiles from Iran. Vidensk Medd Dansk Naturh Foren, 117: 193–207
Shafiei Bafti S., Sehati Sabet M. E., Fahimi H. 2010. A Study on the Reptiles of Southeastern Iran. Kerman, Iran: DOE of Kerman, 220 pp
Sindaco, R., Venchi, A., Grieco, C. 2013. The Reptiles of the Western Palearctic, Volume 2: AnnotatedChecklist and Distributional Atlas of the Snakes of Europe, North Africa, Middle East and Central Asia, with an Update to Volume 1. Latina, Italy: Societas Herpetologica Italica, 543 pp
?míd J., Moravec J., Kodym P., Kratochvíl L., Hosseinian Yousefkhani S. S., Rastegar-Pouyani E., Frynta D. 2014. Annotated checklist and distribution of the lizards of Iran. Zootaxa, 3855: 1–97
Strauch A. A. 1863. Characteristik zweier neuer Eidechsen aus Persien. Bull Acad Imp Sci Saint-Petersbourg, 6: 477–480 (In German)
?míd J, Carranza S., Kratochvíl L., Gvo?ík V., Nasher A. K., Moravec J. 2013. Out of Arabia: A Complex Biogeographic History of Multiple Vicariance and Dispersal Events in the Gecko Genus Hemidactylus (Reptilia: Gekkonidae). PLoS ONE, 8(5): e64018
Solovyeva E. N., Poyarkov N. A., Dunayev E. A., Nazarov R. A., Lebedev V. S., Bannikova A. A. 2014. Phylogenetic Relationships and Subgeneric Taxonomy of Toad-Headed Agamas Phrynocephalus (Reptilia, Squamata, Agamidae) as Determined by Mitochondrial DNA Sequencing. Dokl Biol Sci, 455: 119–124
St?ck M., Günther R., B?hme W. 2001. Progress towards a taxonomic revision of the Asian Bufo viridis group: current status of nominal taxa and unsolved problems (Amphibia: Anura: Bufonidae). Zool Abh Staatl Mus Tierkd Dresden, 51: 253–319
Stümpel N., Joger U. 2009. Recent advances in phylogeny and taxonomy of Near and Middle Eastern Vipers – an update. ZooKeys, 31: 179–191
Torki F. 2010. Beschreibung eines neuen Asaccus (Reptilia: Phyllodactylidae) aus der Provinz Lorestan, Iran. Sauria 32: 3-16 (In German and English)
Torki F. 2011. Description of a new species of Carinatogecko (Squamata: Gekkonidae) from Iran. Salamandra 47: 63–70
Torki F., Ahmadzadeh F., Ilgaz C., Avci A., Kumlutas Y. 2011. Description of four new Asaccus Dixon and Anderson, 1973 (Reptilia: Phyllodactylidae) from Iran and Turkey. Amphibia-Reptilia 32: 185–202
Torki F., Fathinia B., Rostami H. A., Gharzi A., Nazari-Serenjeh F. 2011. Beschreibung eines neuen Asaccus (Sauria: Phyllodactylidae) aus dem Iran. Description of a new Asaccus (Sauria: Phyllodactylidae) from Iran. Sauria, 33: 51–62 (In German and English)
Torki F., Manthey U., Barts M. 2011. Ein neuer Hemidactylus OKEN, 1817 aus der Provinz Lorestan, West-Iran, mit Anmerkungen zu Hemidactylus robustus HEYDEN, 1827 (Reptilia: Squamata: Gekkonidae). Sauria 33: 47–56
Vamberger M., Stuckas H., Ayaz D., Graciá E., Aloufi A. A., Els J., Mazanaeva L. F., Kami H. G., Fritz U. 2012. Conservation genetics and phylogeography of the poorly known Middle Eastern terrapin Mauremys caspica (Testudines: Geoemydidae). Org Divers Evol, Doi: 10.1007/s13127–012–0102–6
Van Dijk P. P., Iversonn J. B., Shaffer H. B., Bour R., Rhodin A. G. J. 2012. Turtles of the world, 2012 Update Annotated Checklist of Taxonomy, Synonymy, Distribution, and Conservation Status. Chel Res Mon, 5: 000,243–000,328
Wall F. 1908. Notes on a Collection of Snakes from Persia. J Bombay Nat Hist Soc, 18: 795–805
Werner F. J. M. 1895. ber eine Sammlung von Reptilien aus Persien, Mesopotamien und Arabien. Verh Zoo -Bot Ges Wien, 45: 13–20 (In German)
Werner F. J. M. 1917. Reptilien aus Persien (Provinz Fars). Gesammelt von Herrn Prof.Andreas. Verh Zoo -Bot Ges Wien, 67: 191–220 (In German)
Werner Y. L. 2006. Retraction of Ptyodactylus goldfuss from the fauna of Iran and its replacement by a new species of Asaccus Dixon and Anderson (Reptilia: Sauria: Gekkonidae). Hamadryad 30: 135–140
Wrobel M. 2004. Elsevier’s Dictionary of Reptiles. London, UK: Elsevier Ltd., 758 pp
Zarudny N. A. 1897. Note sur les Reptiles et Amphibiens de la Perse orientale. Ann Mus Zool Acad Imp Sci St. Pétersbourg, 2: 349–361 (In Russian)
Zarudny N. A. 1902. Marschroute der Expedition der kaiserl. Russ. Geogr. Gesell. In Ost-Persien w?hrend der Jahre 1900 und 1901. Ann Mus Zool Acad Imp Sci St. Pétersbourg, 7: 1–9 (In German)


Last Update: 2016-01-25